Apterygota

The Apterygota are a small subclass of primitive insects.

They are insects which have never had wings at any time in their evolutionary history. Their first known occurrence in the fossil record is in the Rhynie chert of the Devonian period, ~400 million years ago.

They have some other primitive traits (features). The nymphs (younger stages) go through little or no metamorphosis, so they look like the adults. They continue to moult throughout life, with multiple instars (adult stages) after reaching sexual maturity. All other insects have only a single sexually mature adult instar.

Males deposit sperm packages rather than fertilising the female internally.